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Ch.15: Miscellaneous Invaders

Key information on occasional structural invaders: centipedes, millipedes, crickets, sowbugs/pillbugs, earwigs, western conifer-seed bug, box elder bug, and clover mite.

🎯 Top 5 Traps

1
Centipedes have ONE pair of legs per segment with LONG antennae; millipedes have TWO pairs per segment with SHORT antennae. Both belong to the group Myriapoda but in different classes — centipedes in Chilopoda, millipedes in Diplopoda. Centipedes live outside under stones and logs (except the house centipede). Millipedes live in leaf litter and migrate in DRY weather.
2
Sowbugs and pillbugs are CRUSTACEANS — NOT insects. They are small oval LAND crustaceans (class Crustacea, genera Porcellio and Armadillidium) more closely related to crabs, lobsters, and shrimp than to other ground invaders. They feed on decaying vegetable matter and fungi. Control: remove boards on ground, flower pots, flat stones; replace mulch with gravel.
3
Western conifer-seed bug — INSECTICIDES ARE NOT EFFECTIVE INDOORS. Use EXCLUSION + habitat modification only. About 3/4 inch, leaf-footed (family Coreidae), faint white stripes on forewings. When they fly, they BUZZ LIKE BUMBLEBEES and expose ORANGE AND BLACK STRIPES on the abdomen. Do not bite or sting. Lay eggs on PINE NEEDLES; invade structures in late fall and early winter, often basking on south-facing walls before entering.
4
Box elder bug control = REMOVE FEMALE (pod-bearing) BOX ELDER TREES. Clover mite barrier goes on the SUNNY (south) side — NOT the shady side. Box elder bug eggs are laid only on female trees; eliminating the host eliminates the source. Clover mites build up on south sides of buildings where habitat optimum temperature is reached on sunny late fall and early spring days. The 2-foot gravel barrier blocks their migration on the sunny side, with shrubs planted in front to diversify the temperature gradient.
5
Camel/cave crickets are more closely related to KATYDIDS than to crickets — despite the name. They are humpbacked, MOTTLED brown, WINGLESS insects with very long legs and antennae. Often compared to spiders but the resemblance is only superficial. They prefer dark, damp, or cool places (basements, crawl spaces, garages) and seldom cause damage.

🔢 Numbers You Must Know

Number
What It Represents
1 pair / segment
Centipede legs per body segment (with LONG antennae) — class Chilopoda
2 pairs / segment
Millipede legs per body segment (with SHORT antennae) — class Diplopoda
Over 1 inch
House centipede (Scutigera coleoptera) length — runs gracefully on many very long legs
About 1 inch
Field cricket adult length (very dark) and European earwig adult length (dark brown)
About 1/2 inch
Eastern box elder bug (Leptocoris trivittatus) adult length — conspicuous black and red
About 3/4 inch
Western conifer-seed bug (Leptoglossus occidentalis) adult length
Less than 1/16 inch
Clover mite (Bryobia praetiosa) body length — bright to dark red
45 to 75 degrees F
Active temperature range for clover mites; egg and mite development suspends below 45 or above 75 in the ground-level habitat
Above 69 degrees F
Clover mite habitat optimum on sunny late fall and early spring days when general air temperatures are lower (south side of buildings)
2 feet wide
Gravel or gravel-over-plastic barrier strip on the SUNNY side of buildings to stop clover mite migration
20-foot band
Closely mowed lawn band to decrease grass protection and temperature insulation for clover mites
0 males
No male clover mites in the United States — females reproduce parthenogenically

🔀 Easily Confused

Pair / Group
Distinguishing Feature
Centipede vs Millipede
Both Myriapoda (many-segmented). Centipede (Chilopoda): 1 pair of legs per segment; LONG antennae; predator; lives under stones/logs (except house centipede indoors). Millipede (Diplopoda): 2 pairs of legs per segment; SHORT antennae; cylindrical body; lives in leaf litter; migrates in DRY weather; can build up in very large numbers.
Sowbug/pillbug vs Centipede/millipede
Both ground-dwelling many-legged. Sowbug/pillbug: class CRUSTACEA — small oval land crustaceans, related to crabs. Centipede/millipede: group MYRIAPODA — many segments with legs per segment. Crustaceans vs. arthropod centipedes/millipedes.
True crickets vs Camel/cave crickets
True crickets (Gryllidae): related to cockroaches; males "sing" by rubbing wings; field cricket ~1 inch dark with ovipositor for soil egg-laying. Camel/cave cricket: more closely related to KATYDIDS; humpbacked, mottled brown, WINGLESS, very long legs and antennae; dark damp basements/crawl spaces.
Field cricket vs House cricket vs Nemobius
All have CYCLICAL populations and migrate into structures. Field cricket: 1 inch, very dark, most commonly seen; eggs in moist soil of ditches, fields. House cricket: cycles of buildup. Nemobius: very small, dark brown.
Cricket entry conditions
Crickets enter when weeds harden and die and rain is SPARSE — DRY conditions, NOT wet. Move out in massive invasions across parking lots and into malls/buildings. Entry is almost always under doors and through open windows.
Box elder bug vs Western conifer-seed bug
Both gradual-metamorphosis late-season invaders. Box elder bug: ~1/2 inch, conspicuous BLACK AND RED, eggs on female (pod-bearing) box elder trees; nymphs bright red. Western conifer-seed bug: ~3/4 inch, leaf-footed, faint white forewing stripes, eggs on PINE NEEDLES; buzzes like a bumblebee with orange/black abdomen stripes when flying.
Box elder bug control vs Western conifer-seed bug control
Box elder bug: REMOVE FEMALE box elder trees; spray tree trunks/foundations with microencapsulated pesticides during migration. Western conifer-seed bug: insecticides NOT effective indoors — use ONLY exclusion (caulk, weatherstrip, screen, repair soffits) and eliminate nearby host trees; vacuum bugs inside.
Clover mite vs other red mites
Clover mite (Bryobia praetiosa): less than 1/16 inch; bright to dark red; smashes to RED STREAK; FRONT LEGS as long as body that move like antennae (key distinguishing feature). NO males in US — females deposit red eggs in bark crevices and building cracks.
Clover mite barrier: SUNNY vs SHADY side
Goes on the SUNNY (south) side — NOT the shady side. Mites build up where habitat reaches optimum temperature on sunny late fall/early spring days. Barrier components: 2-foot gravel/plastic strip + shrubs planted in front (diversifies temperature gradient) + 20-foot band of closely mowed lawn.
Earwig forceps purpose
Earwig forceps grasp INSECT PREY — they do not bite humans despite scary appearance. Earwigs eat other insects and scavenge garbage and moist plant material. Females TEND THEIR YOUNG (unusual among insects) — guard eggs and care for early-stage nymphs.

🐛 Miscellaneous Invader Quick Reference

Species / Group
ID / Biology
Control Approach
House centipede
Scutigera coleoptera (Chilopoda)
Over 1 inch; many very long legs; 1 pair of legs per segment; long antennae. Predator on tiny insects and spiders. Found in basements + uncontinuously occupied rooms.
Light DUSTING in places where they live. Light residual SPRAY if area is damp. Beneficial — frighten people but cause no damage.
Millipedes
Class Diplopoda
Cylindrical many-segmented; 2 pairs of legs per segment; short antennae. Live outdoors in leaf litter; can build up in very large numbers. Migrate in DRY weather into basements, ground floors, window wells.
Remove leaf litter and compost near foundations. Caulk around door/window facings. Weatherstrip. Residual crack-and-crevice around foundations. BAND/BARRIER application around house if very heavy.
Field crickets
Family Gryllidae
About 1 inch, very dark; gradual metamorphosis. Females inject eggs into moist soil with OVIPOSITOR. Mature July-August. Move out in MASSIVE INVASIONS when weeds harden and rain is sparse. CYCLICAL populations.
Caulk + weatherstrip ground-floor entries. Direct spray into cracks near foundation, door stoops, patios. Residual barrier if populations very high. Granular baits. In high-buildup years: spray weeds and grass in MIDSUMMER with cricket-labeled pesticides.
Camel/cave crickets
Humpbacked, MOTTLED BROWN, WINGLESS; very long legs and antennae. More related to KATYDIDS than crickets. Dark damp basements, crawl spaces, garages. Seldom cause damage.
Ventilate and remove materials providing hiding places. Dusts in crawl spaces and garages may be used but seldom needed. Swat or general-use contact aerosol indoors.
Sowbugs and pillbugs
Class CRUSTACEA — Porcellio + Armadillidium
Small oval LAND CRUSTACEANS (NOT insects). Feed on decaying vegetable matter and fungi. Heavy outside infestations cause individuals to find their way inside.
Habitat alteration only. Remove boards on ground, flower pots, flat stones near house. Remove mulch and replace with gravel if necessary.
European earwig
Forficula auricularia
Dark brown, almost 1 inch; FORCEPS at tail end (grasp insect prey, NOT humans). Wings fold many times under small wing covers. Active at night. Females TEND YOUNG. Hide under bark/boards/stones, in tree wrappings.
Caulk ground-floor entries + cracks between door stoops/patios and foundation. Trim hedges/plants from foundations. Ventilate/dehumidify. Band of low-mowed grass with residual sprays/granules. PESTICIDAL SOAPS (detergents) kill quickly. Dust in dry basements.
Western conifer-seed bug
Leptoglossus occidentalis (Coreidae)
About 3/4 inch; LEAF-FOOTED (hind legs with leaf-like expansions); faint white stripes on forewings. BUZZES LIKE BUMBLEBEE when flying, exposing ORANGE AND BLACK STRIPES on abdomen. Lay eggs on PINE NEEDLES. Late fall/early winter invader; bask on south walls before entering.
INSECTICIDES NOT EFFECTIVE INDOORS — exclusion only. Install/replace weatherstripping; repair/replace screens; caulk around windows/doors; screen vents and chimneys; repair soffits; store firewood outside; eliminate nearby host trees; vacuum bugs inside.
Box elder bug
Leptocoris trivittatus
Conspicuous BLACK AND RED, ~1/2 inch; gradual metamorphosis. Eggs laid in spring on FEMALE (pod-bearing) box elder trees. Young nymphs BRIGHT RED. Mature nymphs/adults crawl down trunk and disperse in late summer; enter wall voids under siding.
BEST METHOD: REMOVE FEMALE (pod-bearing) box elder trees. Caulk entry points on foundation, door, window facings. Spray tree trunks + foundations with microencapsulated pesticides during migration. Vacuum or contact aerosols indoors. Pesticidal soaps.
Clover mite
Bryobia praetiosa
Less than 1/16 inch; bright to dark RED; smashes to RED STREAK. FRONT LEGS as long as body, move like antennae. NO males in US. Females deposit red eggs in bark crevices and building cracks. Active 45-75 degrees F. Build up on SUNNY (south) side of buildings.
OUTSIDE: 2-foot GRAVEL/PLASTIC BARRIER on SUNNY side; plant shrubs in front; 20-foot band of closely mowed lawn; caulk building cracks. INSIDE: caulk window/door framing + electrical plates; cooking oil film on windowsills as temporary trap; vacuum (don't sweep — smears them); spot treatment; crack and crevice; dust voids. SULPHUR is a possible miticide.

💡 Memory Hooks

Centi vs milli: "Centi = ONE pair + LONG antennae; milli = TWO pairs + SHORT." Despite their names, neither has 100 or 1,000 legs — but the per-segment rule is consistent.
Pillbugs are crustaceans: "Pillbugs and sowbugs are crabs in disguise." Class Crustacea — not insects.
Conifer-seed bug: "Caulk and exclude — never spray indoors." Insecticides are not effective.
Box elder bug: "Remove the female trees, remove the bug." Source elimination beats spraying.
Clover mite barrier: "Sunny side, 2 feet of gravel, shrubs in front." South side of buildings, not shady side.
Cave cricket misnomer: "Cave crickets are really katydids." The name is misleading.
Earwig forceps: "Forceps grab insects, not ears." Earwigs prey on other insects; the name and pinchers are misleading about human risk.
Cricket invasion cue: "Dry weeds and sparse rain bring crickets indoors." Wet conditions don't drive cricket invasions; dry conditions do.
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